Adaptive Evolution of Social Traits: Origin, Trajectories, and Correlations of Altruism and Mobility

............................................................................................................................ 3 Introduction ...................................................................................................................... 4 Model Description ............................................................................................................ 6 Population dynamics on social networks..................................................................... 7 Evolutionary dynamics on social networks ................................................................. 8 Spatial invasion fitness .............................................................................................. 10 Selective Pressures on Mobility and Altruism ............................................................... 13 Separate Evolution of Mobility and Altruism ................................................................ 17 Altruism ..................................................................................................................... 17 Mobility ..................................................................................................................... 18 Joint Evolution of Altruism and Mobility ...................................................................... 20 Origin of altruism....................................................................................................... 20 Evolutionary trajectories of social traits .................................................................... 21 Evolutionarily induced correlations between altruism and mobility ......................... 22 Discussion....................................................................................................................... 24 Joint evolution versus single-trait evolution.............................................................. 24 Correlative patterns of social traits ............................................................................ 26 The habitat saturation hypothesis .............................................................................. 29 Concluding remarks................................................................................................... 32 Acknowledgements ........................................................................................................ 33 Appendix A .................................................................................................................... 34 Appendix B..................................................................................................................... 36 Literature Cited............................................................................................................... 39 Table 1 ............................................................................................................................ 45 Figure Legends ............................................................................................................... 46 Figures ............................................................................................................................ 49